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Day 4

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Lecture 4:
Fitnesses that vary in time:

-




Diploid: protected polymorphisms:

- Consider the situation when q is close to 0
- A2 mainly in heterozygotes
- Marginal fitness A2, close to the fitness of heterozygote
- Marginal fitness A1, close to the fitness of A1A1 homozygote.

,Diploid: protected polymorphisms

- Then, analogous to the haploid case, one can show:

,-

, Spatial variation in fitnesses

- Levene’s model:
o Individuals spend their life in a niche/environmental patch
 Own environment
 Own selection
o Emerge from niche only to mate
 At random
 Each niche contributes fixed proportion
o This is soft selection as the extent to which individiuals are eliminated by section in a
niche has no effect on its output
o In contras ot hard selection in which each niche contributes to the mating population
(slide 8)

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